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Beelzebufo ampinga (/biːˌɛlzɪˈbjuːfoʊ æmˈpɪŋɡə/ or /ˌbiːlzəˈbjuːfoʊ/) was a particularly large species of prehistoric frog described in 2008. Common names assigned by the popular media include Killer Bean,[1] devil toad,[2] and the frog from hell.[3]
Fossils of Beelzebufo have been recovered from strata of the Maevarano Formation in Madagascar, dating to the late Cretaceous period, some 70 million years ago (Mya)
The generic name Beelzebufo is a portmanteau of Beelzebub (a Semitic deity whose name may be translated as "Lord of the Flies", sometimes identified either as one of the chief lieutenants, or alter ego of the Christian Devil) and bufo (Latin for "toad").
The specific name ampinga means "shield" in Malagasy.
Animals of this species may have grown to at least 23.2 cm (9.1 in) (snout-vent length), which is still around the size a modern African Bullfrog can reach.[5] The head of Beelzebufo was very big,[5] and bones of the skull roof show a rugous external surface, indicating at least parts of the head may have borne bony scales, called scutes.
The skull sutures are open in even the biggest species of Beelzebufo, showing that it may have grown even bigger. Some estimates suggest snout-vent lengths of up to or beyond 40 cm (15.7 in).[4]
In comparison with the living Ceratophryidae, Beelzebufo most likely was a predator whose expansive mouth allowed it to eat relatively large prey, perhaps even juvenile dinosaurs.[8] Bite force measurements from a growth series of Cranwell's horned frog (Ceratophrys cranwelli), suggest that the bite force of a large Beelzebufo (skull width 15.4 cm) may have been between 500 and 2200 N
The first fossil fragments were found in 1993 by David W. Krause of New York's Stony Brook University, but it took 14 years for scientists Susan E. Evans, Marc E. H. Jones, and Krause to assemble enough data for publication in the Proceedings of the National Academy of Sciences, the journal of the United States National Academy of Sciences.
Some 75 fossil fragments have been found. Researchers have been able to reconstruct parts of the frog's skeleton, including nearly the entire skull.
Horned lizards (Phrynosoma), also known as horny toads or horntoads, are a genus of North American lizards and the type genus of the family Phrynosomatidae. The common names refer directly to their flattened, rounded body and blunt snout.
The genus name Phrynosoma means "toad-bodied". In common with large true frogs and toads, horned lizards tend to move sluggishly, making them easy to catch; this may also avoid triggering attacks by predators. They are adapted to arid or semi-arid areas. The spines on the lizard's back and sides are made from modified reptile scales which prevent the water loss through the skin, whereas the horns on the head are true horns (i.e. they have a bony core). Of the 22 species of horned lizards, 15 are native to the United States. The largest-bodied and most widely distributed of the US species is the Texas horned lizard.
Horned lizards use a wide variety of means to avoid predation. Their coloration generally serves as camouflage. When threatened, their first defense is to remain still to avoid detection. If approached too closely, they generally run in short bursts and stop abruptly to confuse the predator's visual acuity. If this fails, they puff up their bodies to cause them to appear more horned and larger, so that they are more difficult to swallow.[citation needed]
At least eight species (P. asio, P. cornutum, P. coronatum, P. ditmarsi, P. hernandesi, P. orbiculare, P. solare, and P. taurus) are also able to squirt an aimed stream of blood from the corners of the eyes for a distance of up to 5 feet (1.5 m).[1][2][3][4] They do this by restricting the blood flow leaving the head, thereby increasing blood pressure and rupturing tiny vessels around the eyelids. The blood not only confuses predators, but also tastes foul to canine and feline predators. It appears to have no effect against predatory birds. Only three closely related species (P. mcallii, P. modestum, and P. platyrhinos) are certainly known to be unable to squirt blood.[2]
While previous thought held that compounds were added to the blood from glands in the ocular sinus cavity, current research has shown that the chemical compounds that make up the defense are already in the circulating blood.[2][3] It is possible that their diet of large quantities of venomous harvester ants could be a factor; however, the origin and structure of the chemicals responsible are still unknown. The blood-squirting mechanism increases survival after contact with canine predators;[3] therefore, it is probable that, while unorthodox, the trait could have provided an evolutionary advantage. Ocular autohemorrhaging has also been documented in other lizards,[5] which suggests blood-squirting could have evolved from a less extreme defense in the ancestral branch of the genus. Recent phylogenic research supports this claim, so it appears as though the species incapable of squirting blood have lost the adaptation for reasons yet unstudied.[6]
To avoid being picked up by the head or neck, a horned lizard ducks or elevates its head and orients its cranial horns straight up, or back. If a predator tries to take it by the body, the lizard drives that side of its body down into the ground so the predator cannot easily get its lower jaw underneath.[citation needed]
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